Formula: y ~ (((2 * (1 - d) * (1 - (1 - r)^k)) + (2 * d * (1 - (1 - r)^k)^2) + 
    (2 * d * ((1 - r)^k) * (1 - (1 - r)^k))) * dnbinom(x, size = kmercov/bias, 
    mu = kmercov) * length + (((1 - d) * ((1 - r)^k)) + (d * 
    (1 - (1 - r)^k)^2)) * dnbinom(x, size = kmercov * 2/bias, 
    mu = kmercov * 2) * length + (2 * d * ((1 - r)^k) * (1 - 
    (1 - r)^k)) * dnbinom(x, size = kmercov * 3/bias, mu = kmercov * 
    3) * length + (d * (1 - r)^(2 * k)) * dnbinom(x, size = kmercov * 
    4/bias, mu = kmercov * 4) * length)

Parameters:
         Estimate Std. Error t value Pr(>|t|)    
d       2.331e-02  1.285e-03   18.14   <2e-16 ***
r       2.405e-02  3.449e-05  697.37   <2e-16 ***
kmercov 3.861e+01  1.257e-02 3073.06   <2e-16 ***
bias    1.829e+00  5.862e-03  312.07   <2e-16 ***
length  4.758e+06  4.409e+03 1078.96   <2e-16 ***
---
Signif. codes:  0 '***' 0.001 '**' 0.01 '*' 0.05 '.' 0.1 ' ' 1 

Residual standard error: 491.9 on 926 degrees of freedom

Number of iterations to convergence: 7 
Achieved convergence tolerance: 5.015e-07