Formula: y ~ (((2 * (1 - d) * (1 - (1 - r)^k)) + (2 * d * (1 - (1 - r)^k)^2) + 
    (2 * d * ((1 - r)^k) * (1 - (1 - r)^k))) * dnbinom(x, size = kmercov/bias, 
    mu = kmercov) * length + (((1 - d) * ((1 - r)^k)) + (d * 
    (1 - (1 - r)^k)^2)) * dnbinom(x, size = kmercov * 2/bias, 
    mu = kmercov * 2) * length + (2 * d * ((1 - r)^k) * (1 - 
    (1 - r)^k)) * dnbinom(x, size = kmercov * 3/bias, mu = kmercov * 
    3) * length + (d * (1 - r)^(2 * k)) * dnbinom(x, size = kmercov * 
    4/bias, mu = kmercov * 4) * length)

Parameters:
         Estimate Std. Error  t value Pr(>|t|)    
d       7.093e-03  3.669e-04    19.33   <2e-16 ***
r       1.352e-03  6.383e-06   211.89   <2e-16 ***
kmercov 3.699e+01  3.194e-03 11580.59   <2e-16 ***
bias    1.618e+00  2.355e-03   687.00   <2e-16 ***
length  4.436e+06  2.206e+03  2010.68   <2e-16 ***
---
Signif. codes:  0 '***' 0.001 '**' 0.01 '*' 0.05 '.' 0.1 ' ' 1 

Residual standard error: 189.2 on 978 degrees of freedom

Number of iterations to convergence: 6 
Achieved convergence tolerance: 1.339e-06