Formula: y ~ (((2 * (1 - d) * (1 - (1 - r)^k)) + (2 * d * (1 - (1 - r)^k)^2) + 
    (2 * d * ((1 - r)^k) * (1 - (1 - r)^k))) * dnbinom(x, size = kmercov/bias, 
    mu = kmercov) * length + (((1 - d) * ((1 - r)^k)) + (d * 
    (1 - (1 - r)^k)^2)) * dnbinom(x, size = kmercov * 2/bias, 
    mu = kmercov * 2) * length + (2 * d * ((1 - r)^k) * (1 - 
    (1 - r)^k)) * dnbinom(x, size = kmercov * 3/bias, mu = kmercov * 
    3) * length + (d * (1 - r)^(2 * k)) * dnbinom(x, size = kmercov * 
    4/bias, mu = kmercov * 4) * length)

Parameters:
         Estimate Std. Error t value Pr(>|t|)    
d       9.849e-03  1.130e-03    8.72   <2e-16 ***
r       6.462e-03  2.301e-05  280.81   <2e-16 ***
kmercov 2.073e+01  8.471e-03 2447.04   <2e-16 ***
bias    1.525e+00  7.929e-03  192.37   <2e-16 ***
length  1.179e+08  1.326e+05  889.73   <2e-16 ***
---
Signif. codes:  0 '***' 0.001 '**' 0.01 '*' 0.05 '.' 0.1 ' ' 1 

Residual standard error: 15190 on 989 degrees of freedom

Number of iterations to convergence: 8 
Achieved convergence tolerance: 4.034e-06